THE SCIENTIFIC RESEARCH PROGRAM



    Regrowth from decapitation many years prior to uncovering.
     
    Regrowth on Lophophora williamsii -- harvested many years before being uncovered for this photo.



        
      Peyote post-harvest regrowth –
      the harvested group


      Summary of our observations.
      After three years

        PNF – Plant not found.
            TNF – Tag not found.
             nd   – No data available.
               –    – Missing data.


        Concerning data on regrowth involving multiple pups it is important to understand that there is not necessarily a direct 1:1 correspondence between sequential figures for individual pups within a single plant.
         It was not uncommon for some or all pups to be partly or entirely covered so some photos illustrate the pre-uncovery state.

        Images of the tags can be viewed via the links for the tag numbers in the first column and images of the actual plants that were harvested in our study can be viewed using the links in the column headed "Regrowth as number of pups".

        Images of the regrowth plants that were reharvested in the second phase of our study can be viewed using the links in the comments column titled "Reharvested 8 March 2010".

        On 8 March 2010 the second phase of this study began with the two year old regrowth of half the original harvested plants being reharvested. Those twenty plants were rooted for replanting and the mother plants are now part of a subgroup in the study. {The results from our March 2009–2011 visits concerning those 20 individuals.}




      Plant No.
       [Tag  IMG]
      Number
      of ribs

       
      Diameter
      (cm)
       
      Height
      (cm)
       
      Weight
      (gm)
       
      Regrowth as
      number of pups
      or TNF, PNF
      Regrowth as
      diameter of pups

      (cm)
         Comments   

       
      2008
      2009
      2010
      2011
              3 Mar. 2008 (This refers to the original plant.)


       Also entries for 2010-11
      11 Nov. 2008
        7 Mar. 2009
       6 Mar. 2010
      15 Mar. 2011
       11 Nov. 2008
         7 Mar. 2009
        6 Mar. 2010
      15 Mar. 2011
       
       
      101 
      101
      101
      101
      13
      –
      8, 7, 6, 7
       nd
      6.2


       
      3.0


       
      65


       
      4
      4
      4
      0
      1.3, 2.5, 2.4, 1.2
      2.0, 2.7, 2.5, 1.3
      2.5, 3.1, 3.1, 20
      nd


        Reharvested 8 March 2010. 
        Dead remains found. 
      102 
      102
      102
      102
        5
      –
      nd
      nd
      2.4


       
      1.0


       
        3


       
      1
      1
      nd
      0
      1.5
      1.5
      nd
      nd


        Plant not present. Hog activity. 
        Plant not present.
      103 
      103
      103
      103
      5
      –
      7, 7
      6, 6
      2.0


       
      0.7


       
        3


       
      3
      2
      2
      2
      0.5, 0.8, 0.9
      1.2, 1.3
      1.4, 1.5
      0.6, 0.6

      104 
      104
      104
      104
        8
      –
      *
      nd
      4.6


       
      1.5


       
      23


       
      TNF, PNF
      TNF, PNF
      2
      0
      nd
      nd
        9
      nd


        *Not countable. 
        Carcass present. 
      105 
      105
      105
      105
        8
      –
      nd
      nd
      5.0


       
      1.5


       
      23


       
      0
      TNF, PNF
      0
      0
      nd
      nd
      nd
      nd


        Presumed dead but not locateable.
       
      106 
      106
      106
      106
      13
      –
      7, 5
      not countable
      6.3


       
      1.6


       
      45


       
      2
      3
      2
      1
      2.0, 2.4
      2.0, 2.4, 1.9
      4.0, 7.0
      1.0



        Partially herbivored. 
      107 
      107
      107
      107
        8
      –
      nd
      nd
      5.0


       
      3.0


       
      31


       
      6
      6
      0
      0
      1.5, 2.0, 1.8, 1.3, 1.6, 1.4
      1.8, 2.4, 2.2, 1.5, 1.9, 1.5
      nd
      nd


         Dead. Remains located. 
       Abundant remains still present
      108 
      108
      108
      108
        8
      –
      5, 7, 7
      7, 7, 5*
      5.0


       
      1.5


       
      31


       
      TNF, PNF
      3
      3
      3
      nd
      2.1, 1.8, 2.1
      2.1, 1.6, 2.4
      2.1, 1.3, 1.3


        Reharvested 8 March 2010.
        *Chewed on by herbivores. 
      109 
      109
      109
      109
        9
      –
      7, 8, 7
      8, 7, 7
      5.8


       
      1.9


       
      43


       
      TNF, PNF
      TNF, PNF
      3
      3
      nd
      nd
      2.9, 3.8, 3.4
      3.1, 3.0, 3.1

      110 
      110
      110
      110
        8
      –
      7, 7
      nd
      4.1


       
      1.8


       
      23


       
      TNF, PNF
      TNF, PNF
      2
      0
      nd
      nd
      2.5, 2.1
      nd


        Reharvested 8 March 2010.
        Dead. Carcass located. 
      111 
      111
      111
      111
        8
      –
      8, 5
      6
      2.7


       
      0.9


       
        6


       
      TNF, PNF
      TNF, PNF
      2
      1
      nd
      nd
      2.0, 0.6
      1.0

      112 
      112
      112
      112
      13
      –
      8, 7, 8, 8
      7, 8, 6, 6
      4.8


       
      0.7


       
      20


       
      TNF, PNF
      4
      4
      4
      nd
      2.1, 2.0, 1.7, 1.6
      2.6, 1.3, 2.7, 2.4
      1.3, 1.5, 1.3, 1.3


        Reharvested 8 March 2010.
       
       113 
      113
      113
      113
      13
      –
      nd
      nd
      6.3


       
      2.6


       
      74


       
      1
      0
      0
      0
      0.9
      nd
      nd
      nd


          Dead. Remains located.
        Carcass still present.
      114 
      114
      114
      114
        8
      –
      7, 8
      8, 5
      4.1


       
      1.2


       
      17


       
      2
      2
      2
      2
      1.7, 1.9
      1.7, 1.7
      2.1, 2.2
      2.3, 1.0


        Reharvested 8 March 2010.
        1 head looks older than the other.
      115 
      115
      115
      115
        5
      –
      nd
      nd
      2.2


       
      0.8


       
        3


       
      0
      0
      0
      0
      nd
      nd
      nd
      nd


        Dead but not locateable. 
        Carcass found. 
      116 
      116
      116
      116
        8
      –
      8, 5, 6
      5*, 8, 6
      4.4


       
      0.3


       
      23


       
      3
      3
      3
      3
      1.7, 1.0, 1.4
      1.7, 0.9, 1.6
      2.5, 0.7, 1.8
      2.5, 2.5, 0.7



        *Chewed on by herbivores. 
      117 
      117
      117
      117
        8
      –
      11, 6, 6
      8, 7, 12
      5.4


      1.7


      28


      3
      3
      3
      3
      1.3, 2.8, 1.8
      1.8, 3.0, 2.1
      1.7, 3.6, 2.2
      2.6, 2.5, 4.5

      118 
      118
      118
      118
        8
      –
      8, 7, 7, 6, 6
      7, 7, 6
      5.3


       
      1.3


       
      34


       
      5
      5
      5
      3
      1.0, 1.3, 1.0, 1.2, 1.4
      1.0, 1.9, 1.3, 1.7, 2.0
      1.9, 1.1, 1.4, 1.0, 1.5
      1.7, 1.8, 1.8


        Reharvested 8 March 2010.
       
      119 
      119
      119
      119
      13
      –
      8, 5, 8, 8
      6, 7, 7, 6
      5.4


       
      2.3


       
      48


       
      4
      4
      4
      4
      1.9, 0.8, 1.5, 1.6
      2.2, 0.9, 1.3, 1.9
      2.4, 0.9, 1.6, 1.6
      2.4, 2.8, 3.2, 1.9

      120 
      120
      120
      120
        8
      –
      7, 8
      7, 6
      5.6


       
      1.4


       
      28


       
      TNF, PNF
      TNF, PNF
      2
      2
      nd
      nd
      1.8, 2.1
      2.0, 1.5


        Reharvested 8 March 2010.
        One crown was stepped on. 
      121 
      121
      121
      121
      13
      –
      8, 7
      7, 7
      6.3


       
      2.6


       
      71


       
      2
      2
      2
      2
      1.9, 2.5
      2.4, 2.6
      2.4, 2.7
      3.6, 3.4

      122 
      122
      122
      122
        5
      –
      7
      6
      2.9


       
      1.0


       
        6


       
      1
      TNF, PNF
      1
      1
      2.2
      nd
      2.1
      1.7

        Reharvested 8 March 2010.
      123 
      123
      123
      123
        8
      –
      6, 5
      7, 6
      5.0


       
      1.8


       
      31


       
      TNF, PNF
      TNF, PNF
      2
      2
      nd
      nd
      2.1, 1.6
      2.5*, 2.9



        *Regrowth after herbivore damage. 
      124 
      124
      124
      124
        8
      –
      7, 6
      5, 5
      4.1


       
      1.1


       
      17


       
      2
      2
      2
      2
      1.9, 2.1
      1.8, 2.0
      1.8, 1.8
      0.8, 2.0


        Reharvested 8 March 2010.
       
      125 
      125
      125
      125
        8
      –
      5, 6
      6, 7
      4.4


       
      1.1


       
      14


       
      2
      2
      2
      2
      1.6, 1.5
      2.0, 1.6
      1.4, 1.4
      2.5, 2.5

      126 
      126
      126
      126
      13
      –
      6, 7, 7
      7, 6, 8, 7
      5.0


       
      2.0


       
      54


       
      TNF, PNF
      TNF, PNF
      3
      4
      nd
      nd
      2.4, 2.5, 2.4
      1.8, 1.9, 2.0, 1.4


        Reharvested 8 March 2010.
       
      127 
      127
      127
      127
        8
      –
      8
      8
      4.3


       
      1.2


       
      28


       
      1
      1
      1
      1
      1.8
      2.0
      2.3
      3.0

      128 
      128
      128
      128
      13
      –
      6, 8, 8
      6, 7, 6, 7
      4.9


       
      1.6


       
      34


       
      3
      3
      3
      4
      1.6, 2.4, 2.0
      1.7, 2.8, 2.0
      1.8, 2.8, 2.1
      1.5, 1.7, 1.6, 1.5


        Reharvested 8 March 2010.
       
      129 
      129
      129
      129
      10
      –
      6, 7, 7
      8, 8, 7, 7
      5.3


       
      2.0


       
      34


       
      4
      4
      3
      4
      2.0, 2.0, 1.0, 2.2
      2.2, 2.2, 1.7, 2.3
      2.2, 2.0, 2.3
      2.8, 2.8, 2.8, 2.8

      130 
      130
      130
      130
        8
      –
      7, 7
      13, 8
      5.3


       
      1.6


       
      23


       
      2
      2
      2
      2
      1.1, 3.1
      1.1, 3.7
      3.9, 1.4
      4.7, 2.3

      131 
      131
      131
      131
        8
      –
      7, 7, 7
      6
      4.8


       
      2.6


       
      26


       
      3
      3
      3
      1
      1.4, 1.2, 1.1
      1.9, 1.5, 1.4
      1.0, 1.7, 1.6
      2.0


        Reharvested 8 March 2010.
        Lopsided; herbivores ate one side. 
      132 
      132
      132
      132
        8
      –
      nd
      nd
      4.4


       
      1.5


       
      17



      2
      2
      nd
      0
      1.5, 2.0
      1.9, 2.0
      nd
      nd


        Plant not present. Hog activity. 
       
      133 
      133
      133
      133
        8
      –
      6
      6
      4.2


       
      1.7


       
      20


       
      0
      1
      1
      1
      nd
      0.8
      1.3
      2.0

      134 
      134
      134
      134
        8
      –
      6, 7
      nd
      3.4


       
      1.1


       
        9


       
      0
      2
      2
      0
      nd
      1.4, 1.6
      1.4, 1.4
      nd


        Reharvested 8 March 2010.
       
      135 
      135
      135
      135
      13
      –
      12, 5
      12, 6
      5.4


       
      1.0


       
      28


       
      2
      2
      2
      2
      2.8, 1.7
      3.2, 1.9
      3.9, 1.8
      4.5, 2.4

      136 
      136
      136
      136
      13
      –
      6, 7, 7
      5*, 6, 6
      5.7


       
      2.3


       
      48


       
      2
      3
      3
      3
      1.9, 1.9
      2.4, 2.3, 1.4
      2.1, 2.2, 2.0
      1.2, 1.7, 1.3


        Reharvested 8 March 2010.
        *Apex of plant off-center. 
      137 
      137
      137
      137
        8
      –
      6, 6, 6
      6, 7, 7
      4.6


       
      1.5


       
      17


       
      3
      3
      3
      3
      1.8, 1.8, 1.7
      1.6, 1.6, 1.3
      1.3, 1.2, 1.2
      2.1, 2.0, 2.0

      138 
      138
      138
      138
      13
      –
      6, 5, 5
      6
      5.7


       
      2.6


       
      43


       
      TNF, 4
      4
      3
      1
      1.1, 1.4, 1.8, 1.1
      1.2, 1.7, 2.4, 1.4
      3.0, 1.2, 1.2
      1.9


        Reharvested 8 March 2010.
        Apical meristem eaten and regrown.
      139 
      139
      139
      139
        5
      –
      6
      8
      2.9


       
      1.0


       
        6


       
      1
      1
      1
      1
      1.8
      1.9
      2.1
      2.6

      140 
      140
      140
      140
      10
      –
      nd
      nd
      5.4


       
      1.2


       
      28


       
      0
      0
      0
      0
      nd
      nd
      nd
      nd


          Presumed dead but not locateable. 
       
      141 
      141
      141
      141
      13
      –
      7, 7, 7, 7
      8, 9, 5*
      5.0


       
      1.0


       
      23


       
      4
      4
      4
      3
      1.6, 1.6, 1.8, 1.5
      –, –, –, –
      2.0, 1.8, 1.5, 1.5
      1.8, 2.0, 1.5


        Reharvested 8 March 2010.
        *Pup still in the process of emerging.
      142 
      142
      142
      142
      13
      –
      6, 6, 6, 6
      8, 7, 7, 6
      6.4


       
      2.1


       
      62


       
      TNF, PNF
      TNF, PNF
      4
      4
      nd
      nd
      2.6, 1.3, 1.1, 1.2
      2.0, 2.0, 2.0, 3.3

      143 
      143
      143
      143
      13
      –
      8, 8, 7
      7, 9, 7
      6.3


       
      1.9


       
      51


       
      TNF, PNF
      TNF, PNF
      3
      3
      nd
      nd
      2.9, 2.7, 2.6
      1.7, 2.8, 2.1

        Reharvested 8 March 2010.
      144 
      144
      144
      144
        5
      –
      6, 6
      6, 6
      3.9


       
      1.7


       
      23


       
      2
      2
      2
      2
      1.0, 1.0
      1.4, 1.3
      0.9, 1.0
      2.0, 2.0

      145 
      145
      145
      145
        5
      –
      5
      7
      3.5


       
      0.9


       
      11


       
      1
      1
      1
      1
      1.4
      1.5
      1.4
      1.6


        Reharvested 8 March 2010.
       
      146 
      146
      146
      146
      13
      –
      7, 8
      8, 9
      5.9


       
      1.5


       
      37


       
      2
      2
      2
      2
      1.8, 1.6
      2.0, 1.9
      2.2, 2.3
      2.8, 2.8

      147 
      147
      147
      147
      13
      –
      7, 7, 7, 6
      8, 8, 8
      5.7


       
      1.6


       
      34


       
      4
      4
      4
      3
      1.0, 1.4, 1.4, 1.8
      1.5, 1.5, 2.0, 2.0
      1.7, 1.5, 1.7, 1.2
      1.1, 1.5, 1.8


        Reharvested 8 March 2010.
       
      148 
      148
      148
      148
      13
      –
      7
      7
      6.8


       
      3.0


       
      54


       
      1
      1
      1
      1
      2.5
      3.4
      3.2
      4.1

      149 
      149
      149
      149
      13
      –
      8x3, 7x3
      7, 8*
      6.7


      1.5


      51


      6
      6
      6
      2
      1.8, 1.1, 1.0, 0.9, 1.1, 1.8,
      2.1, 2.0, 1.9, 1.4, 2.0, 2.0
      2.9, 1.7, 1.9, 1.9, 1.7, 2.4
      2.3, 2.4


        Reharvested 8 March 2010. 
        *Deformed. 
      150 
      150
      150
      150
      5
      –
      6, 7
      8, 7
      2.9


       
      1.0


       
        9


       
      2
      2
      2
      2
      1.3, 1.3
      1.4, 1.4
      1.0, 1.2
      2.3, 2.3


      The plants being surveyed are part of a larger population occurring on private land in South Texas. The area of the survey site itself is defined with a transect of 17-gauge electric fence wire secured with steel stakes — the locations of which are recorded with a high resolution GPS. The transect zigzags through the study site with all plants in the study being located within approximately one-half meter of the wire. The area of the study site has not yet been accurately measured but we estimate it to be around half a hectare.

          We also tagged and measured a control group of 50 plants on 23 NOV 2008, which in future monitoring will give us an idea of the magnitude of natural mortality not associated with harvesting. {The results from our visits concerning those 50 individuals serving as our control group.}

          An additional 20 plants (#201-220) were harvested during the November 2008 monitoring to make up for what was believed to be lost plants from the original group of 50 harvested plants. {The results from our visits concerning those twenty individuals.}


      7 March 2009 update:
          Interestingly we found the same number of harvested study plants (with their tags) but as can be seen in the table above these were not entirely the same plants & tags as had been located on 23 November 2008.
          On this visit we used a Trimble high-resolution GPS instrument to record the location of all tags and/or plants located.
          Complete or partial burial of plants and their tags was sometimes encountered. To aid in future visits we added a rebar stake near each tagged plant.


      6 March 2010 update:
          A number of our tags and plants were completely buried and hence provided missing data in 2009. In 2010, thanks to the use of a high powered metal detector, we located all of our tags, stakes and plants unless the plants themselves were totally missing from the study site.




          Feral hog activity was extensive at this site so we will split the presentation of our final data assessment into two conclusions: mortality involving hogs and mortality not involving hogs.
      Lophophora is surviving many known challenges.





      Regrowth study site showing typical Tamaulipan thornscrub vegetation

         

          If you would like to help support additional studies of this type, designed to answer basic questions that are critical to the optimal stewardship/management of vulnerable cactus species, please send a contribution, of $25 USD or whatever amount you can afford, to the Cactus Conservation Institute, 909 E. Sul Ross Ave., Alpine, TX 79830.
          We also now accept donations via PayPal.






    Please enjoy our featured articles:


    A Tale of Two Cacti


    by Dana M. Price & Martin Terry



    Button, button, who's got the button?


    by Martin Terry


    Conservation of Lophophora in Mexico


    by Martin Terry

    Flora of the Dead Horse Mountains

    by Joselyn Fenstermacher






    All images © are copyrighted by the photographers and/or the CCI and are used with permission.
    Reproduction is forbidden without prior written consent.




    Cactus Conservation Institute